Dr. Baroni

Dr. Baroni

Timothy J. Baroni,

Distinguished Professor

Boletus aurantiosplendens
Boletus aurantiosplendens Baroni
-- a new species from North Carolina

& Lab:
340 Bowers Hall
Phone: 607-753-2725
E-mail: tim.baroni@cortland.edu


Humboldt State University, B.A., M.A.
University of Massachusetts, Amherst, Ph.D.

Courses Frequently Taught

  • Biological Sciences I  (BIO 201)
  • Plants and People (SCI 304)
  • Fungi
  • Field Mycology
  • Scanning Electron Microscopy

Scholarly Interests

Studies on the systematics and biodiversity of mushrooms, especially members of the Entolomataceae, and boletes (Boletaceae) are the focal point for much of my research.  Phylogenetic analyses of members of the Entolomataceae, in collaboration with Drs. Valerie Hofstetter and Jean-Marc Moncalvo, using several different molecules, have played a prominent role in my recent efforts, in addition to broad based field work and morphological investigations, in an effort to develop a more complete understanding of this family of mushrooms on a world wide scale. 

In the past 10 years most of my time has been spent collecting and studying tropical mushrooms and boletes from the islands of Puerto Rico, Dominican Republic and Jamaica in the Greater Antilles as well as Belize and Costa Rica in Central America. Through several major collaborative efforts we have made good progress in documenting the biodiversity of Basidiomycetes from these regions.  Our group (for more information visit:  www.cortland.edu/nsf/ga.html) has catalogued over 7,000 collections so far from the Greater Antilles project.  Numerous presentations at national and international meetings and a growing list of technical papers have been published on the results of our work.  We have described many new species of basidiomycetes (and have many more yet to be described), we have also described some new genera (see papers below), and we believe we may have discovered at least one new family of basidiomycetes which is yet to hit the press.  Numerous new records have been recorded for the Greater Antilles and Belize, and we are beginning to develop ideas about the biogeography of basidiomycetes for this region of the Neotropics. 

Recent opportunities have allowed me to study first hand Asian and Australasian mushrooms after making a trip to Thailand and Tasmania to work with colleagues.  We (Genevieve Gates and I) have published one paper on new species of Rhodocybe from Tasmania, and other papers are planned on some members of the Tricholomataceae from Tasmania.  Work on Asian collections started some years ago through collaborations with Drs. Roy Watling and Dennis Desjardin. My recent visit to this part of the world has provided me with a new prospective and lots of my own collections to examine.  

Cortland Undergraduate Biology Students continue to work with me on research projects and Mr. Lance Lacey (class of 2004) and Ms. Nicole Bocsusis (class of 2007) have been instrumental in preparing publications on new species and new reports.  Lance joined our team on several research expeditions to the Dominican Republic and Belize from 2001 to 2004 and made major contributions to documenting collections from those countries.

Recent Grants

    • NSF - “ Basidiomycetes of the Greater Antilles, Especially the Luquillo LTER site”   $452,643.00   co-pi with Drs. D. Jean Lodge, Karen Nakasone and Leif Ryvarden (1996-2000)
    • Faculty Research Program (Cortland):  “Molecular taxonomy in the Entolomataceae”   $2,317.50
    • NSF – Supplement award to the Basidiomycetes of the Greater Antilles grant to run a second workshop on “Tropical Basidiomycete Taxonomy and Systematics” in Venezuela, August 1999,  $22,000.00, co-pi with Dr. D. Jean Lodge.
    • Sponsored Programs (Cortland) Incentive Grant – molecular systematics/phylogenetics of Clitopilus (Entolomataceae), DNA sequencing of the ITS region of rDNA at Duke University, summer 2000,  $1,000.00
    • NSF – “Basidiomycetes of Neotropical Pine Forests: Connections Between a  Possible Refugium in Belize and Endemic Pine Forests in the Dominican Republic.” co-PI with Drs. D. Jean Lodge, L. Ryvarden, J. Cifuentes, and O. K. Miller, Jr. $399,981. (2001-2005)


  • NSF – REU award to support two undergraduate students (Cortland and University of Puerto Rico) on Basidiomycetes of Neotropical Pine Forests, $12,000
  • National Science Foundation – supplement award to the Basidiomycetes of Neotropical Pine Forests:  to invite young investigators from Costa Rica, Venezuela, Colombia, Germany, France and the US to present their research findings on Diversity of Neotropical Macrofungi in a symposium organized by me and Dr. Roy E. Halling, New York Botanical Garden, at the 7th International Mycological Congress in Oslo, Norway, 11-17 August 2002, $11,000.00
  • National Geographic Society -– Mycological biodiversity of Doyle’s Delight, Maya Mts. Belize, $11,725 (Aug. 2007)
  • British Mycological Society -– Mycological biodiversity of Doyle’s Delight, Maya Mts., Belize,$975 (Aug. 2007)

Selected Publications

Note that the underlined publications below have links to pdf files.

  • Ryvarden, L., Aime, M. C., and Baroni, T. J. 2009. Studies in neotropical polypores 26. A new species of Trametes and revisitation of an old. SYNOPSIS FUNGORUM FUNGIFLORA 26:27-32. 
  • Largent, D.L., Henkel, T.W., Aime, M. C., and Baroni, T.J. 2008. The Entolomataceae of the Pakaraima Mountains of Guyana I: four new species of Entoloma ss.str. MYCOLOGIA 100: 132-140.
  • Baroni, T. J., Bocsusis, N., Lodge, D. J., Lindner, D. L.  2008. A new species of Pleurocollybia (Tricholomataceae; Agaricales; Basidiomycetes) from Belize. MYCOTAXON (in press)
  • Baroni, T. J., Franco-Molano, A. E., Lodge, D. L., Lindner, D. L., Hofstetter, V., and Horak, E. 2007. Arthromyces and Blastosporella two new genera of lyophylloid agarics (Agaricales, Basidiomycota) from the Neotropics that produce conidia.  MYCOLOGICAL RESEARCH 111:572-580.

  • Petersen, R. H. and Baroni, T. J. 2007. Xerula hispida and Xerula setulosa (comb. nov.) two similar subtropical New World agarics MYCOTAXON 101:113-136.

  • Halling, R.E., Baroni, T. J., and Binder, M. 2007.  A New Genus of Boletaceae from Eastern North America. MYCOLOGIA 99:310-316.

  • Ovrebo, C. L. and Baroni, T. J. 2007. New taxa of Tricholomataceae and Entolomataceae (Agaricales) from Central America. Fungal Diversity 27-157-170.

  • Ortiz-Santana, B., Lodge, D. J., Baroni, T. J, and Both, E. E. 2007. Boletes from Belize and the Dominican Republic. Fungal Diversity 27:247-416

  • Matheny, P. Brandon, Zheng Wang, Manfred Binder, Judd M. Curtis, Young Woon Lim, R. Henrik Nilsson, Karen W. Hughes, Valerie Hofstetter,Joseph F. Ammirati, Conrad L. Schoch, Ewald Langer, Gitta Langer, David J. McLaughlin, Andrew W. Wilson, Tobias Frøslev, Zai-Wei Ge, Richard W. Kerrigan, Jason C. Slot, Zhu-Liang Yang, Timothy J. Baroni, Michael Fischer, Kentaro Hosaka, Kenji Matsuura, Michelle T. Seidl, Jukka Vauras, David S. Hibbett. 2007.  Contributions of rpb2 and tef1 to the phylogeny of Mushrooms and allies (Basidiomycota, Fungi). Molecular Phylogenetics and Evolution 43: 430–451.

  • Matheny, P. Brandon, Judd M. Curtis, Valerie Hofstetter, M. Catherine Aime, Jean-Marc Moncalvo, Zai-Wei Ge, Zhu-Liang Yang, Jason C. Slot Joseph F. Ammirati, Timothy J. Baroni, Neale L. Bougher, Karen W. Hughes, D. Jean Lodge, Richard W. Kerrigan, Michelle T. Seidl, Duur K. Aanen, Matthew DeNitis, Graciela M. Daniele, Dennis E. Desjardin, Bradley R. Kropp, Lorelei L. Norvell, Andrew Parker, Else C. Vellinga, Rytas Vilgalys,David S. Hibbett. 2006 .Major clades of Agaricales: a multilocus phylogenetic overview. Mycologia, 98(6):982–995.

  • Baroni, T. J. and Gates, G. 2006. New species and records of Rhodocybe (Entolomataceae, Agaricales) from Tasmania.  Australian Systematic Botany 19: 343-358.

  • Armstrong, T. R. and Baroni, T. J.  2004. A new species of Mycena (Tricholomataceae, Agaricales) from the central mountains of the Dominican Republic.  In Fungi in Forest Ecosystems – Systematics, Diversity, and Ecology.  Dedicated to Dr. Orson K. Miller, Jr. Cathy L. Cripps, editor.  Memoirs of the New York Botanical Garden 89:125-129. 2004.

  • Moncalvo, J-M, Baroni, T. J., Bhatt, R. P., Stephenson, S. S.  2003. Rhodocybe paurii, a new species from the Indian Himalaya.  Mycologia 96:859-865.

  • Guzmán, G., Tapia, F., Guillén, F. R., Baroni, T. J., Lodge, D. J., Cantrell, S. A., and Nieves-Rivera, A. M.  2003. New species of Psilocybe in the Caribbean with an emendation of P. guilartensis. Mycologia 95:1171-1180.

  • Lodge, D.J., T.J. Baroni, and S.A. Cantrell. 2002. Basidiomycetes of the Greater Antilles project. pp. 45-60 In: R. Watling, J. C. Frankland, A. M. Ainsworth, S. Isaac & C. Robinson, eds., Tropical Mycology, Vol. 1, Macromycetes. CABI Publishing Egham, UK.

  • Moncalvo, J.M., R. Vilgalys, S.A. Redhead, J.E. Johnson, T.Y. James, M.C. Aime, V. Hofstetter, S.J.W. Verduin, E. Larsson, T.J. Baroni, R.G. Thorn, S. Jacobsson, H. Clémençon, and O.K. Miller Jr. 2002. One Hundred and Seventeen Clades of Euagarics. Molecular Phylogenetics and Evolution 23:357-400.

  • Aime, M. C., Baroni, T. J. and Miller, O. K., Jr.  2002. Crepidotus thermophilus comb. nov., a reassessment of Melanomphalia thermophila, a rarely collected tropical agaric. MYCOLOGIA 94:1059-1065

  • Baroni, T. J. and Ortiz, B.  2002. New species of Oudemansiella and Pouzarella (Basidiomycetes: Agaricales) from Puerto Rico.  MYCOTAXON 82:269-279

  • Baroni, T. J. and Mattock, G.  2001. Rhodocybe roseiavellanea (Murr.) Singer, a new record for Britain and Europe. THE MYCOLOGIST 15(4):151-152.

  • Cantrell, S. A., Lodge, D. J., and Baroni, T. J.  2001. Basidiomycetes of the Greater Antilles Project. THE MYCOLOGIST 15(3):107-112.

  • Baroni, T. J.,  Desjardin, D. E., and Hywel-Jones, N.  2001. Clitopilus chalybescens, a new species from Thailand.  FUNGAL DIVERSITY 6:13-17.

  • Miller, O. K., Jr., D. J. Lodge, and T. J. Baroni.  2000. New and interesting ectomycorrhizal fungi from Puerto Rico, Mona, and Guana Islands.  MYCOLOGIA 92(3):558-570.

  • Baroni, T. J. and R. E. Halling.  2000.  Some Entolomataceae (Agaricales) from Costa Rica.  BRITTONIA 52:121-135.

  • Orson K. Miller, Jr., D. J. Lodge, and T. J. Baroni.  2000.  New and interesting ectomycorrhizal fungi from Puerto Rico, Mona, and Guana Islands.  MYCOLOGIA 92:558-570.

  • Baroni, T. J. and R. Watling.  1999.  Taxonomic and mycogeographic notes on some Malaysian fungi IV.  Notes on Clitopilus and Rhodocybe.  MYCOTAXON 72:57-72.